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Identification of Chrysididae

Once prepared, the specimen can be parked in an entomological box, with the similar ones, but one last component it still lacking: the identification label. Such label is written by its identifier, the entomologist who recognizes it on the basis of its diagnostic characters and assigns it to a known species. Therefore, the last pinned label should report the following info: Genus species Author, sex, identifier and year, i.e. Chrysis ignita L., ♀, C.Darwin det., 1980). When more determination cards are present on the same spacimen it means that the specimen has been examined and revised by many specialists, each one leaving his own taxonomic opinion. Generally, the last temporal diagnosis is the most reliable, reported on the lowest label.

Some chrysidids

OcularsThe procedure of identification of a specimen is executed by a specialist who receives the material by a collector. After some time, he returns the material and keeps some specimens for his own collection, as an "honorarium" for his performance. The identification is performed using a stereoscopic microscope and through a critical comparison between what is seen and what is written in the determination keys.

Móczár (1997) wrote: « Most species of Cleptes [but this could be extended to the whole Chrysididae] are rare and their colour and sculpture show a great variety even within populations. The main reasons behind it are the cleptoparasitic lifestyle and the microclimatic circumstances which influence individual development. Thus, their classification is sometimes uncertain. »

The main determination keys for Chrysididae have been supplied by the following Authors: W. Linsenmaier, L. Móczár, L.S. Kimsey and R. Bohart (see the Literature page).

Chrysis.net offers a


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All text and images of this page are copyright ©️ Chrysis.net unless otherwise stated - please see individual cases for authorship and copyright details. The specimens pictured are from the authors' or other collaborators' personal collections and from the collections of various museums. Unless otherwise specified, the whole content of this web site is for personal, non-commercial, scientific, and educational purposes given proper accreditation to the page from which they were derived are provided, and under Chrysis.net Terms and Conditions.

For citation purposes

Agnoli G.L. & Rosa P. (2025) Search Results , in: Chrysis.net website. Interim version 29 April 2025, URL: https://www.chrysis.net/search/%252525E7%25252583%252525AD%252525E8%252525A1%25252580%252525E9%252525AB%25252598%252525E6%252525A0%252525A11~3%252525E7%25252599%252525BE%252525E5%252525BA%252525A6%252525E7%252525BD%25252591%252525E7%2525259B%25252598-%252525E3%25252580%25252590%252525E2%2525259C%25252594%252525EF%252525B8%2525258F%252525E6%2525258E%252525A8%252525E8%2525258D%25252590BB76%252525C2%252525B7CC%252525E2%2525259C%25252594%252525EF%252525B8%2525258F%252525E3%25252580%25252591-%252525E5%25252585%252525AB%252525E4%252525B8%252525AA%252525E8%2525259D%252525B4%252525E8%2525259D%252525B6%252525E5%25252585%252525AC%252525E4%252525B8%252525BB%252525E6%25252595%25252585%252525E4%252525BA%2525258B-%252525E7%25252583%252525AD%252525E8%252525A1%25252580%252525E9%252525AB%25252598%252525E6%252525A0%252525A11~3%252525E7%25252599%252525BE%252525E5%252525BA%252525A6%252525E7%252525BD%25252591%252525E7%2525259B%25252598nvq0t-%252525E3%25252580%25252590%252525E2%2525259C%25252594%252525EF%252525B8%2525258F%252525E6%2525258E%252525A8%252525E8%2525258D%25252590BB76%252525C2%252525B7CC%252525E2%2525259C%25252594%252525EF%252525B8%2525258F%252525E3%25252580%25252591-%252525E5%25252585%252525AB%252525E4%252525B8%252525AA%252525E8%2525259D%252525B4%252525E8%2525259D%252525B6%252525E5%25252585%252525AC%252525E4%252525B8%252525BB%252525E6%25252595%25252585%252525E4%252525BA%2525258Bw2t0-%252525E7%25252583%252525AD%252525E8%252525A1%25252580%252525E9%252525AB%25252598%252525E6%252525A0%252525A11~3%252525E7%25252599%252525BE%252525E5%252525BA%252525A6%252525E7%252525BD%25252591%252525E7%2525259B%25252598yut6s-%252525E5%25252585%252525AB%252525E4%252525B8%252525AA%252525E8%2525259D%252525B4%252525E8%2525259D%252525B6%252525E5%25252585%252525AC%252525E4%252525B8%252525BB%252525E6%25252595%25252585%252525E4%252525BA%2525258B3oas/page/17.

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Taxonomy of the Hymenoptera

Hymenoptera is a large order of insects, comprising over 150,000 described species. They originated in the Triassic, and social hymenopterans during the Cretaceous.

The Classification of the Hymenoptera consists of two main divisions (starting from Gerstäcker, 1867):

  1. the Symphyta, which lack a "wasp waist", and instead have a broad connection between the abdomen (metasoma) and the thorax (mesosoma);
  2. the Apocrita, which have a narrow waist "wasp waist".

Symphyta - The suborder Symphyta includes the sawflies, horntails, and parasitic wood wasps. The Symphyta are the more primitive (basal) group of the Hymenoptera, with an unconstricted junction between the thorax and abdomen and comparatively complete wing venation. The larvae are phytophagous (herbivorous), with three pairs of true legs, prolegs (on every segment, unlike Lepidoptera) and ocelli. Together, the Symphyta make up less than 10% of hymenopteran species, with about 10,000 described species.

Apocrita - The suborder Apocrita includes the true wasps, bees, and ants. The Apocrita are the most advanced hymenopterans. They have a constriction between the first and second abdominal (metasomal) segments called a "wasp-waist" (petiole), with the first abdominal segment fused to the thorax (propodeum). The larvae of all Apocrita lack legs, prolegs, or ocelli and are blind. The family Orussidae (Symphyta) may be the group from which the Apocrita arose.

The Apocrita have historically been split into two groups, "Parasitica" and "Aculeata", but these are rankless groupings.

Parasitica – The term Parasitica is an artificial (paraphyletic) group comprising the majority of hymenopteran insects, with members living as parasitoids, many of which having an ovipositor adapted for piercing.

Aculeata – The Aculeata are a monophyletic group, or good natural group, representing a single "branch" on the "tree of life". It includes those species in which the female's ovipositor is modified into a stinger to inject venom. The Aculeata are therefore maintained as infraorder or division or as an unranked clade – From Wikipedia (https://en.wikipedia.org/wiki/Aculeata).

Taxonomy of the Hymenoptera

The taxonomy of Hymenoptera can be based on different classification criteria, such as morphology, behavior, cladistic methods and molecular phylogenetics. Here we present three alternative hierarchical trees.

This tree does not consider the phylogenetic relationships, since many different visions are still discordant. And is a taxonomy based on Superfamilies that abandons the traditional approach of using sections (infraorders) as ‘Terebrantia’ (‘Parasitica’) or ‘Aculeata’ of the Suborder ‘Apocrita’.

 

Kingdom Animalia

Subkingdom Eumetazoa

Phylum Arthropoda

Subphylum Hexapoda

Class Insecta

Order Hymenoptera

Family Gerocynipidae
Family Palaeocynipidae
Family Rasnitsyniidae

Suborder Symphyta

Superfamily Cephoidea
Family Cephidae
Superfamily Megalodontoidea
Family Megalodontidae
Family Pamphilidae
Superfamily Orussoidea
Family Orussidae
Superfamily Siricoidea
Family Siricidae
Superfamily Tenthredinoidea
Family Argidae
Family Blasticotomidae
Family Cimbicidae
Family Diprionidae
Family Pergidae
Family Tenthredinidae
Superfamily Xyeloidea
Family Anaxyelidae
Family Xyelidae
Family Xiphydriidae

Suborder Apocrita

Superfamily Apoidea
Family Ampulicidae
Family Andrenidae
Family Apidae
Family Colletidae
Family Crabronidae
Family Halictidae
Family Heterogynaidae
Family Megachilidae
Family Melittidae
Family Sphecidae
Family Stenotritidae
Superfamily Ceraphronoidea
Family Ceraphronidae
Family Megaspilidae
Superfamily Chalcidoidea
Family Agaonidae
Family Aphelinidae
Family Chalcididae
Family Eucharitidae
Family Eulophidae
Family Eupelmidae
Family Eurytomidae
Family Leucospidae
Family Mymaridae
Family Ormyridae
Family Perilampidae
Family Pteromalidae
Family Rotoitidae
Family Signiphoridae
Family Tanaostigmatidae
Family Tetracampidae
Family Torymidae
Family Trichogrammatidae
Superfamily Chrysidoidea
Family Bethylidae
Family Chrysididae
Family Dryinidae
Family Embolemidae
Family Plumariidae
Family Sclerogibbidae
Family Scolebythidae
Superfamily Cynipoidea
Family Austrocynipidae
Family Cynipidae
Family Figitidae
Family Ibaliidae
Family Liopteridae
Superfamily Evanioidea
Family Aulacidae
Family Evaniidae
Family Gasteruptiidae
Superfamily Ichneumonoidea
Family Braconidae
Family Ichneumonidae
Superfamily Megalyroidea
Family Megalyridae
Superfamily Mymarommatoidea
Family Mymarommatidae
Superfamily Platygastroidea
Family Platygastridae
Family Scelionidae
Superfamily Proctotrupoidea
Family Austroniidae
Family Diapriidae
Family Heloridae
Family Monomachidae
Family Pelecinidae
Family Peradeniidae
Family Proctorenyxidae
Family Proctotrupidae
Family Roproniidae
Family Vanhorniidae
Superfamily Stephanoidea
Family Stephanidae
Superfamily Trigonalyoidea
Family Trigonalyidae
Superfamily Vespoidea
Family Bradynobaenidae
Family Formicidae
Family Mutillidae
Family Pompilidae
Family Rhopalosomatidae
Family Sapygidae
Family Scoliidae
Family Sierolomorphidae
Family Tiphiidae
Family Vespidae

 

 

 

 

Source: https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=152741#null

Kingdom Animalia — animals

Subkingdom Bilateria

Infrakingdom Protostomia

Superphylum Ecdysozoa

Phylum Arthropoda - arthropods

Subphylum Hexapoda — hexapods

Class Insecta — insects

Subclass Pterygota — winged insects

Infraclass Neoptera — modern, wing-folding insects

Superorder Holometabola

Order Hymenoptera - ants, bees, wasps

Suborder Apocrita - wasps, bees, and ants

Infraorder Aculeata - "stinging wasps"

Superfamily Apoidea - bees, sphecoid wasps, apoid wasps
Family Ampulicidae - cockroach wasps, ampulicid wasps
Family Andrenidae - andrenid bees, andrenids
Family Angarosphecidae
Family Apidae - bumble bees, euglossine, euglossines, honey bees, stingless bees
Family Colletidae - colletid bees, plasterer bees, yellow-faced bees
Family Crabronidae - crabronid wasps, cicadakillers, sand wasps, mud daubers
Family Halictidae - halictid bees, sweat bees
Family Heterogynaidae
Family Megachilidae - leafcutting bees
Family Melittidae - melittid bees, melittids
Family Paleomelittidae
Family Sphecidae - mud daubers, sphecid wasps, digger wasps, sand wasps
Family Stenotritidae - stenotritid bees, stenotritids
Superfamily Bethylonymoidea
Family Bethylonymidae
Superfamily Chrysidoidea
Family Bethylidae - bethylid wasps
Family Chrysididae - cuckoo wasps
Family Dryinidae - dryinid wasps
Family Embolemidae
Family Plumariidae
Family Sclerogibbidae
Family Scolebythidae
Superfamily Vespoidea - vespoid wasps
Family Bradynobaenidae
Family Falsiformicidae
Family Formicidae - ants, fourmis
Family Mutillidae - velvet ants
Family Pompilidae - spider wasp
Family Rhopalosomatidae - rhopalosomatid wasps
Family Sapygidae - sapygid wasps
Family Scoliidae - scoliid wasps
Family Sierolomorphidae - sierolomorphid wasps
Family Tiphiidae - tiphiid wasps
Family Vespidae - hornets, paper wasps, potter wasps, yellowjackets

Infraorder Terebrantes

Superfamily Ceraphronoidea
Family Ceraphronidae - ceraphronids, ceraphronides, ceraphronid wasps
Family Maimetshidae
Family Megaspilidae - megaspilides, megaspilid wasps, megaspilids
Family Stigmaphronidae
Superfamily Chalcidoidea - chalcidoid wasps, chalcidoids
Family Agaonidae - fig wasps
Family Aphelinidae
Family Chalcididae - chalcidid wasps, chalcids, chalcidids
Family Encyrtidae - encyrtid wasps, encyrtids
Family Eucharitidae - eucharitids
Family Eulophidae - eulophid wasps, eulophids
Family Eupelmidae - eupelmids
Family Eurytomidae - eurytomids, jointworms, seed chalcids
Family Leucospidae - leucospids
Family Mymaridae - fairyflies
Family Ormyridae - armyrids
Family Perilampidae - perilampids
Family Pteromalidae - pteromalid wasps, pteromalids
Family Rotoitidae
Family Signiphoridae
Family Tanaostigmatidae - tanaostigmatids
Family Tetracampidae
Family Torymidae - torymid wasps, torymids
Family Trichogrammatidae - trichogrammatid wasps, trichogrammatids
Superfamily Cynipoidea - gall wasps, cynipoids
Family Archaeocynipidae
Family Austrocynipidae
Family Cynipidae - gall wasps, cynipid gall wasps
Family Figitidae - figitids
Family Ibaliidae - ibaliids
Family Liopteridae - liopterids
Family Rasnicynipidae
Superfamily Evanioidea - parasitic hymenoptera
Family Aulacidae - aulacids
Family Evaniidae - ensign wasps
Family Gasteruptiidae - gasteruptiids
Family Praeaulacidae
Superfamily Ichneumonoidea - parasitic hymenoptera, hyménoptères parasites
Family Braconidae - braconid wasps, braconids, braconides
Family Eoichneumonidae
Family Ichneumonidae - ichneumon flies, ichneumon wasps, ichneumons
Family Praeichneumonidae
Superfamily Megalyroidea
Family Megalyridae
Superfamily Mymarommatoidea
Family Mymarommatidae
Superfamily Platygastroidea
Family Platygastridae - platygastrids
Family Scelionidae - scelionid wasps, scelionids
Superfamily Proctotrupoidea - true parasitic hymenoptera
Family Austroniidae
Family Diapriidae - diapriids
Family Heloridae - helorids
Family Jurapriidae
Family Maamingidae
Family Mesoserphidae
Family Monomachidae
Family Pelecinidae - pelecinid wasps, pelecinids
Family Peradeniidae
Family Proctorenyxidae
Family Proctotrupidae - proctotrupids
Family Roproniidae - roproniids
Family Vanhorniidae - vanhorniids
Superfamily Serphitoidea
Family Serphitidae
Superfamily Stephanoidea
Family Stephanidae - stephanids
Superfamily Trigonaloidea
Family Trigonalidae

Suborder Symphyta - sawflies, horntails, and parasitic wood wasps

Superfamily Cephoidea - stem sawflies
Family Cephidae - stem sawflies
Superfamily Orussoidea
Family Orussidae - parasitic wood wasps
Superfamily Pamphilioidea
Family Megalodontesidae
Family Pamphiliidae - leaf-rolling sawflies, web-spinning sawflies
Superfamily Siricoidea - wood wasps
Family Anaxyelidae
Family Siricidae - horntails
Superfamily Tenthredinoidea - sawflies
Family Argidae - argid flies, argid sawflies
Family Blasticotomidae
Family Cimbicidae - cimbicid sawflies
Family Diprionidae - conifer sawflies
Family Pergidae - pergid flies
Family Tenthredinidae - common sawflies, tenthredinid sawflies
Superfamily Xiphydrioidea
Family Xiphydriidae - wood wasps
Superfamily Xyeloidea
Family Xyelidae - xyelid flies, xyelid sawflies

Recent research in molecular phylogenetics has resulted in the reorganization of the infraorder Aculeata, which now contains eight superfamilies: Apoidea, Chrysidoidea, Formicoidea, Pompiloidea, Scolioidea, Tiphioidea, Thynnoidea, and Vespoidea.

 
NOTE: the symbol † indicates an extinct taxon.
 

Kingdom Animalia – animals

Subkingdom Bilateria

Infrakingdom Protostomia

Superphylum Ecdysozoa

Phylum Arthropoda – arthropods

Subphylum Hexapoda – hexapods

Class Insecta – insects

Subclass Pterygota – winged insects

Infraclass Neoptera – modern, wing-folding insects

Superorder Holometabola

Order Hymenoptera – ants, bees, wasps

Suborder Apocrita – wasps, bees, and ants

(unranked) Aculeata - "stinging wasps"

Superfamily Apoidea subgroup Spheciformes (previously: "Sphecoidea") – sphecoid wasps
Family Ammoplanidae
Family Ampulicidae (cockroach wasps)
Family Astatidae
Family Bembicidae
Family Crabronidae (sand wasps, bee wolves, etc.)
Family Mellinidae
Family Pemphredonidae
Family Philanthidae
Family Psenidae
Family Sphecidae (digger wasps)
Superfamily Apoidea subgroup Anthophila (previously: "Apoidea") – bees, apoid wasps
Family Andrenidae (mining bees)
Family Apidae (carpenter bees, digger b., cuckoo b., bumble b., orchid b., and honeybees)
Family Colletidae (yellow-faced bees and plasterer bees)
Family Halictidae ("sweat bees")
Family Megachilidae (leaf-cutting bees)
Family Melittidae
Family Stenotritidae
Superfamily Chrysidoidea
Family Bethylidae
Family Chrysididae (cuckoo wasps)
Family Dryinidae
Family Embolemidae
Family Plumariidae
Family Sclerogibbidae
Family Scolebythidae
Superfamily Formicoidea
Family Formicidae (ants)
Superfamily Pompiloidea
Family Mutillidae (velvet ants)
Family Myrmosidae
Family Pompilidae (spider wasps)
Family Sapygidae
Superfamily Scolioidea
Family Scoliidae
Superfamily Tiphioidea
Family Bradynobaenidae
Family Sierolomorphidae
Family Tiphiidae
Superfamily Thynnoidea
Family Chyphotidae
Family Thynnidae
Superfamily Vespoidea
Family Rhopalosomatidae
Family Vespidae (paper wasps, potter wasps, hornets, pollen wasps, yellowjackets)

(unranked) Parasitica

Superfamily Ceraphronoidea
Family Ceraphronidae
Family Megaspilidae
Superfamily Chalcidoidea
Family Agaonidae (fig wasps)
Family Aphelinidae
Family Chalcididae (chalcid wasps)
Family Encyrtidae
Family Eucharitidae
Family Eulophidae
Family Eupelmidae
Family Eurytomidae (seed chalcids)
Family Leucospidae
Family Mymaridae (fairyflies) — the smallest of all insects
Family Ormyridae
Family Perilampidae
Family Pteromalidae
Family Rotoitidae
Family Signiphoridae
Family Tanaostigmatidae
Family Tetracampidae
Family Torymidae
Family Trichogrammatidae
Superfamily Cynipoidea
Family Austrocynipidae
Family Cynipidae (gall wasps)
Family Figitidae
Family Ibaliidae
Family Liopteridae
Superfamily Diaprioidea
Family Austroniidae
Family Diapriidae
Family Maamingidae
Family Monomachidae
Superfamily Evanioidea
Family Aulacidae
Family Evaniidae (ensign wasps)
Family Gasteruptiidae
Superfamily Ichneumonoidea
Family Braconidae
Family Ichneumonidae (ichneumon wasps)
Superfamily Megalyroidea
Family Megalyridae
Superfamily Mymarommatoidea
Family Mymarommatidae
Superfamily Platygastroidea
Family Platygastridae
Family Scelionidae
Superfamily Proctotrupoidea
Family Heloridae
Family Pelecinidae
Family Peradeniidae
Family Proctorenyxidae
Family Proctotrupidae
Family Roproniidae
Family Vanhorniidae
Superfamily Stephanoidea
Family Stephanidae
Superfamily Trigonaloidea
Family Trigonalidae

Suborder Symphyta - sawflies, horntails, and parasitic wood wasps

Superfamily Anaxyeloidea
Family Anaxyelidae
Superfamily Cephoidea
Family Cephidae
Family †Sepulcidae
Superfamily †Karatavitoidea
Family †Karatavitidae
Superfamily Orussoidea
Family Orussidae
Superfamily Pamphilioidea
Family Megalodontesidae
Family Pamphiliidae Cameron, 1890
Family †Xyelydidae
Superfamily Siricoidea
Family Siricidae (horntails)
Family Xiphydriidae (wood wasps, sometimes treated as a separateSuperfamily, Xiphydrioidea)
Family †Daohugoidae
Family †Protosiricidae
Family †Pseudosiricidae
Family †Sinosiricidae
Superfamily Tenthredinoidea
Family Argidae
Family Blasticotomidae
Family Cimbicidae
Family Diprionidae
Family Pergidae
Family Tenthredinidae
Superfamily Xiphydrioidea
Family Xiphydriidae
Superfamily Xyeloidea
Family Xyelidae

References

Aguiar, A.P.; Deans, A.R.; Engel, M.S.; Forshage, M.; et al. (2013). "Order Hymenoptera. In: Zhang Z-Q (ed) Animal biodiversity: An outline of higher-level classif. and survey of taxonomic richness". Zootaxa. 3703: 51–62.

Debevec A.H., Cardinal S., Danforth B.N. (2012) "Identifying the sister group to the bees: a molecular phylogeny of Aculeata with an emphasis on the superfamily Apoidea". Zoologica Scripta 41: 527-535.

Pilgrim, Erik M.; Von Dohlen, Carol D.; Pitts, James P. (2008). "Molecular phylogenetics of Vespoidea indicate paraphyly of the superfamily and novel relationships of its component families and subfamilies". Zoologica Scripta. 37 (5): 539–560.

Sharkey, M.J.; Carpenter, J.M.; Vilhelmsen, L. (2012). "Phylogenetic relationships among superfamilies of Hymenoptera". Cladistics. 28: 80–112.


Copyright, Authorship, and Ownership statements

All text and images of this page are copyright ©️ Chrysis.net unless otherwise stated - please see individual cases for authorship and copyright details. The specimens pictured are from the authors' or other collaborators' personal collections and from the collections of various museums. Unless otherwise specified, the whole content of this web site is for personal, non-commercial, scientific, and educational purposes given proper accreditation to the page from which they were derived are provided, and under Chrysis.net Terms and Conditions.

For citation purposes

Agnoli G.L. & Rosa P. (2025) Search Results , in: Chrysis.net website. Interim version 29 April 2025, URL: https://www.chrysis.net/search/%252525E7%25252583%252525AD%252525E8%252525A1%25252580%252525E9%252525AB%25252598%252525E6%252525A0%252525A11~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~3%252525E7%25252599%252525BE%252525E5%252525BA%252525A6%252525E7%252525BD%25252591%252525E7%2525259B%25252598nvq0t-%252525E3%25252580%25252590%252525E2%2525259C%25252594%252525EF%252525B8%2525258F%252525E6%2525258E%252525A8%252525E8%2525258D%25252590BB76%252525C2%252525B7CC%252525E2%2525259C%25252594%252525EF%252525B8%2525258F%252525E3%25252580%25252591-%252525E5%25252585%252525AB%252525E4%252525B8%252525AA%252525E8%2525259D%252525B4%252525E8%2525259D%252525B6%252525E5%25252585%252525AC%252525E4%252525B8%252525BB%252525E6%25252595%25252585%252525E4%252525BA%2525258Bw2t0-%252525E7%25252583%252525AD%252525E8%252525A1%25252580%252525E9%252525AB%25252598%252525E6%252525A0%252525A11~3%252525E7%25252599%252525BE%252525E5%252525BA%252525A6%252525E7%252525BD%25252591%252525E7%2525259B%25252598yut6s-%252525E5%25252585%252525AB%252525E4%252525B8%252525AA%252525E8%2525259D%252525B4%252525E8%2525259D%252525B6%252525E5%25252585%252525AC%252525E4%252525B8%252525BB%252525E6%25252595%25252585%252525E4%252525BA%2525258B3oas/page/17.

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Systematics of Chrysididae

The fundamental morphologic features which distinguish chrysidids from the other wasps are:

  • reduction of the number of the visible abdominal segments to 5 or less
  • an internal ovipositor or genital tube, constituted by the remaining abdominal segments
  • 11 antennal articles (flagellomeres), plus pedicel and scape
  • labial palps with 3 articles and maxillar palps with 5 articles
  • fore wings with 5 closed cells, or less, and
  • hind wings without closed cells.

According to the Zoological Record, Chrysidids are ranked in the taxonomic hierarchy as Aculeate Wasps, wasps equipped by a stinging apparatus. But, according to the present-day classifications, the traditional sections (infraorders) 'Terebrantia' ('Parasitica') and 'Aculeata' of the Suborder 'Apocrita' have been abandoned in the direction of taxonomy based on Superfamilies.

The Family Chrysididae comprises 5 Subfamilies and 83 Genuses. Their taxonomy down to the Genus level is as follows:

  • Class Insecta
    • Order Hymenoptera
      • Suborder Apocrita
        • Series Aculeata (obsolete)
          • Superfamily Chrysidoidea
            • Family Chrysididae
              • Subfamily Cleptinae
                • Cleptes*
                • Cleptidea
              • Subfamily Amiseginae
                • Adelphe
                • Afrosega
                • Alieniscus
                • Amisega
                • Anachrysis
                • Anadelphe
                • Atoposega
                • Baeosega
                • Bupon
                • Cladobethylus
                • Colocar
                • Duckeia
                • Exopapua
                • Exova
                • Imasega
                • Indothrix
                • Isegama
                • Kimseya
                • Kryptosega
                • Leptosega
                • Magdalium
                • Mahinda
                • Microsega
                • Myrmecomimesis
                • Nesogyne
                • Nipponosega
                • Obenbergerella
                • Perissosega
                • Reidia
                • Rohweria
                • Saltasega
                • Serendibula
              • Subfamily Loboscelidiinae
                • Loboscelidia
                • Rhadinoscelidia
              • Subfamily Chrysidinae
                • Tribe Allocoeliini
                  • Allocoelia
                • Tribe Elampini
                  • Adelopyga
                  • Chrysellampus*
                  • Colpopyga*
                  • Diplorrhos
                  • Elampus*
                  • Exallopyga
                  • Haba
                  • Hedychreides
                  • Hedychridium*
                  • Hedychrum*
                  • Holophris
                  • Holopyga*
                  • Microchridium
                  • Minymischa
                  • Muesebeckidium
                  • Oligogaster
                  • Omalus*
                  • Parachrum
                  • Philoctetes*
                  • Prochridium
                  • Pseudolopyga
                  • Pseudomalus*
                  • Xerochrum
                • Tribe Kimseyini
                  • Kimseya
                • Tribe Chrysidini
                  • Allochrysis
                  • Argochrysis
                  • Caenochrysis
                  • Ceratochrysis
                  • Chrysidea
                  • Chrysis*
                  • Chrysura*
                  • Chrysurissa
                  • Euchroeus*
                  • Exochrysis
                  • Gaullea
                  • Ipsiura
                  • Istiochrysis
                  • Morphochrysis*
                  • Neochrysis
                  • Odontochrydium
                  • Pentachrysis*
                  • Pleurochrysis
                  • Praestochrysis*
                  • Primeuchroeus
                  • Pseudochrysis*
                  • Spinolia*
                  • Spintharina*
                  • Spintharosoma
                  • Stilbichrysis
                  • Stilbum*
                  • Trichrysis*
              • Subfamily Parnopinae
                • Cephaloparnops
                • Isadelphia
                • Parnopes*

* the asterisk indicates the presence of the Genus in Europe.

 


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All text and images of this page are copyright ©️ Chrysis.net unless otherwise stated - please see individual cases for authorship and copyright details. The specimens pictured are from the authors' or other collaborators' personal collections and from the collections of various museums. Unless otherwise specified, the whole content of this web site is for personal, non-commercial, scientific, and educational purposes given proper accreditation to the page from which they were derived are provided, and under Chrysis.net Terms and Conditions.

For citation purposes

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Field search of Chrysididae

The Chrysidid collection can be carried out on the field through traps (Malaise and yellow dishes) or through sight-hunting with a butterfly net (aluminum handle, nylon net bag).

Malaise trapMalaise trap. This trap consists of a kind of Canadian tent, dark in color, with a vertical wall in the middle which prevents the flying wasps to pass through by forcing them to fly upward. On the corner top of the tent there's a hole which lets the daily light filter through and attracts the wasps which are flying in the direction of the light in order to escape. The hole brings a transparent plastic tube which ends into a large bottle containing a preserving liquid (75% alcohol + water), where the wasps will fall. Malaise trap is cumbersome and needs to be set on the flying routes of the wasps. For this reason, the trap is advisable for areas where it won't attract the attention of too curious people. You need to control the collecting bottle every week, depending on many factors. However, Malaise trap is a massive collecting method, which won't sample only the wasps you are looking for, but everything is flying there, unconditionally.

Moericke trapsMoericke traps (or "yellow pan traps"). You simply need the ordinary plastic dishes used for pic-nic already colored or painted of intense yellow color (we suggest the spray acrylic enamel) and positioned close to the flying routes of the Chrysidids: rocks, paths, wood-piles, seasoned bushes. Once positioned, the plates need to be filled up with a mixture of three elements: water (in order to make the wasp drown), soap (in order to break off the surficial tension of the water and preventing the floating of the wasp) and salt (in order to preserve the specimens for some days). Salt is a good preserving mineral, but it will make the colors of the wasps "opaque" in case of long-lasting permanence; therfore it is important to control the content of the dishes periodically. The meaning of the yellow color comes from the fact that many flying insects orient toward the flowers through visual indications. The yellow color is one of the best colors to attract insects. Yellow is also the wavelength reflected by young leaves.

Sight-hunting. Flowers, rocks, walls, woods, wood-piles, hollow branches, sandy soils inhabited by other wasps/bees, dry walls, palissades, wood stakes, etc. Chrysidids rest generally for few second on these places and then they fly quickly to the next place, usually following an invisible route, probably chemically marked and generally constant. It is not difficult to scare a specimen and, remaining in wait, see it rest again after few minutes on the same point or on the same path. You need to pay attention to your position regarding the sun and the shadow when you approach the resting place of the wasps. Sometimes, when you shot your net on a specimen resting on a wall or on wood, you will see the wasp remain motionless or point to escape under the edge of the net; in these cases you have to "convince" the wasp to fly upwards in the bag. Once the wasp is in the bag, you have to push it in your bottle containing the mortal agent, carrying the bottle inside the bag. The liquid generally used to kill the chrysidids - and all the other insects - is the ethylacetate (EtOAc). We generally use a glass or poliethylene (PP) killing jar with an hermethic cap and a cotton bottom containing some drops of etylacetate (its vapors kills the specimens). Ethyl acetate is toxic when used, so avoid ingestion, inhalation and contact. As an alternative to ethylacetate you can use ammonia (NH3).

NestAn alternative collection method consists in the breeding of nests made by other wasps, from which chrysidids could emerge as hosts. Examples of nests could be the mud cells of Sphecids and of Eumenids, or the Cynipid galls housed by secondary hosts (Sphecids of the genus Pemphredon), or the hollow and dried branches used by many Sphecids and Apoids. Consequently this method allows you to reach a deeper knowledge on Chrysidid relationships with hosts.


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Coloration of Chrysididae

With some exceptions, Chrysidids are generally colored and their structural coloration is due to light interference, and it varies with the viewing angle.

Many species are characterized by colors with metallic glares, green, blue, copper, gold, etc., and some colorations seem to be typical of precise geographic regions.

Chrysis grohmanni krkiana

Stilbum cyanurumThe Loboscelidiini and the Allocoeliini lack any metallic coloration; their colors are brown, black, reddish and, in the some cases, white.

The Amisegini are generally from brown to black, with little metallic glares from green to blue on face and thorax; the abdomen is generally non-metallic (metallic in Duckeia).

The coloration of the Cleptini is very variable. Some species are completely non-metallic black. Cleptes species are generally metallic on head and thorax, the abdomen being non-metallic. Typical of almost all the European Cleptes is the metallic color of the head and of the thorax (mainly red in females and green-blue in males), with a non- or not completely metallic abdomen.

The Chrysidini and the Elampini are always colored with metallic colors. In Europe the commonest colors are the green-blue on head and thorax, with a copper or golden abdomen. The species of southern Spain, N Africa and Middle East tend to be completely copper- or brass-colored. In tropical Asia we observe the diffusion of a pattern with green body and a copper-colored spot on both sides of the second abdominal tergite in species belonging to even distant Genera (the reason is unknown), while in the Philippines Chrysidids are purple with a red-shining head. Some Hedychridium show a reddish non-metallic abdomen.

In Parnopes a clean chromatic distinction based on the geographic distribution is observed. The African and American species tend to be colored of uniform blue, green, purple. In the palearctic species, instead, the abdomen is often different from the rest of the body and generally non-metallic.

White colorations are generally reduced to spots and stripes on mandibles, antennal articles, tegulae, legs, abdominal tergites; in the species of the Loboscelidiini and the Elampini they are not observed.

From: Kimsey L.S. & Bohart R.M., 1991 – The Chrysidid wasps of the World. Oxford University Press, ix-652.

Observations:

  • color can be altered from the chemicals used in order to kill, to preserve or to rehydrate specimens.
  • color has a diagnostic value in many cases, but not always, because each species shows a variability both chromatic and morphologic.
  • does an environmental relation between color of the adult and the physical parameters (humidity, temperature) at the time of the development exists?
  • melanism is rarely observed.

 

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Hosts of Chrysididae

Here you get a short survey on the Hymenoptera Genera parasitized by Chrysidids. The list is mainly based on the data collected for the European fauna. You can take a look at the Hymenoptera hierarchy according to the Vol.134 of the Zoological Record.

The drawings are taken from Yeo P.F. & Corbet S.A., 1983 - Solitary Wasps - Naturalist's Handbook 3, Cambridge, and from Chinery M., 1986 - Collins Guide to the Insects of Britain and Western Europe - Collins.

Detailed data about the hosts of the italian Chrysidids are in the Database of the Italian Chrysididae.

Order Hymenoptera

Suborder Symphyta

Superfamily Tenthredinoidea

Family Tenthredinidae: Caliroa, Nematus, Pristiphora

Tenthredinoidea

Suborder Apocrita

Superfamily Sphecoidea

Family Sphecidae: Astata, Bembix, Cerceris, Crabro, Dinetus, Diodontus, Ectemnius, Harpactus, Larra, Lindenius, Mimesa, Miscophus, Nitela, Oxybelus, Passaloecus, Pemphredon, Philanthus, Pison, Podalonia, Psen, Psenulus, Rhopalum, Solierella, Stigmus, Tachysphex, Tracheliodes, Trypoxylon

Sphecoidea

Superfamily Apoidea

Family Anthophoridae: Anthophora, Eucera
Family Apidae: Ceratina
Family Colletidae: Colletes, Hylaeus
Family Halictidae: Halictus
Family Megachilidae: Anthidium, Anthocopa, Chalicodoma, Chelostoma, Heriades, Hoplitis, Megachile, Osmia, Prosopis, Protosmia, Pseudoanthidium, Rhodanthidium, Trianthidium

Apoidea

Sometimes it happens that Chrysidids emerge from abandoned galls, i.e. the wood galls caused by Cynipid wasps on oak branches. The presence of Chrysidids testifies that Megachilid bees used the cynipid gall as a prefabricate house. The parasitic interaction between Chrysidids and Megachilids does not involve the Cynipids.

Gall

Superfamily Vespoidea

Family Eumenidae: Alastor, Allodynerus, Ancistrocerus, Delta, Discoelius, Eumenes, Euodynerus, Gymnomerus, Microdynerus, Odynerus, Paragymnomerus, Spinicoxa, Symmorphus, Tropidodynerus
Family Masaridae: Celonites, Ceramius
Family Vespidae: Paravespa, Vespula

Vespoidea


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